Unidad del IGME de Zaragoza

Enlaces de interés

Enlaces de interés

Samuel Zamora Iranzo
Doctor en Ciencias Geológicas
Contratado Ramón y Cajal
Actividad: Paleontología
Teléfono: + 34 976 555 153
E-mail: s.zamora@igme.es


Samuel Zamora Iranzo



I graduated in Geology from Zaragoza University in 2004 and obtained my doctorate from the same University in 2009 (funded by the Government of Aragón). My doctorate thesis entitle “Middle Cambrian echinoderms from the Iberian Chains and Cantabrian Zone (North Spain)”, was awarded the maximum score possible (Cum Laude) and won the Best Science Thesis prize from Faculty of Sciences (Zaragoza University) in 2010. I next spent 24 months as a postdoctoral researcher at the Natural History Museum (London) funded by the Spanish Ministry of Science and Education. My proposal “Tracing the origins of pentamery: fossil echinoderm stem groups and their importance for understanding the derivation of major new body plans”, obtained the maximum score in Earth Sciences. My research in London focused on deciphering the early evolution of a major deuterostome group (echinoderms) in order to reveal the detailed pathways that gave rise to new body plans. My research was based on original field collections from various parts of the world (Spain, France, Italy, UK, Check Republic, Morocco, China, USA) and the fossils obtained were studied using a combination of traditional and state-of-the-art imaging techniques. By combining data from phylogenies, three-dimensional imaging, morphometrics and sequence stratigraphy I was able to document how key echinoderm lineages evolved and responded at critical times of global change, namely the Cambrian Explosion and the Great Diversification Ordovician Event. I spent later another PostDoc as a researcher at the Smithsonian Institution at the National Museum of Natural History (Washington DC), a position won in a public and competitive call. My duties were to develop an independent research project using the collections there, and I was pursuing a project entitled “The Early evolution of echinoderms and the timing of clade origination: A comparison between Gondwana and Laurentia”. I am currently employed at the Spanish Geological Survey (IGME) with a Ramón y Cajal Research Grant funded by the Spanish Ministry of Economy and Competitiveness. My work will be focus on Palaeozoic echinoderms from Spain.



- Álvaro, J. J., Zamora, S. & Vizcaïno, D. & Ahlberg, P. 2013.  Guzhangian (mid Cambrian) trilobites from silica concretions of the Valtorres Formation, Iberian Chains, NE Spain. Geological Magazine. 150 (1): 123-142.
- Álvaro, J. J., Zamora, S., Clausen, S., Vizcaïno, D. & Smith, A. B. 2013. Cambrian substrate revolution or evolution? A review from the benthic community replacements of West Gondwana. Earth Science Reviews. 118: 69-82.
- Esteve, J., Hughes, N. & Zamora, S. 2013. Thoracic structure and enrolment style in middle Cambrian Eccaparadoxides pradoanus presages caudalization of the derived trilobite trunk. Palaeontology. 56-3, 589-601.
- Gorzelak, P. & Zamora, S. 2013. Stereom microstructures of Cambrian echinoderms revealed by cathodoluminescence (CL). Paleontologia Electronica.16 (3). 32A.
- Kammer, T., Sumrall, C. D., Zamora, S., Ausich, W. A., & Deline, B. 2013. Oral Region Homologies in Paleozoic Crinoids and other Plesiomorphic Pentaradial Echinoderms. PlosOne 8 (11).
- Smith, A. B., Reich, M., & Zamora, S. 2013. Comment on supposed holothurian body fossils from the middle Ordovician of Wales (Botting and Muir, Palaeoelectronica: 15.1.9A). Paleontologia Electronica. 16.1.3A.
- Smith, A. B., Reich, M. & Zamora, S. Devonian echinoids from Spain and Germany. Acta Palaeontologica Polonica. 58 (4), 751-762.
- Smith, A. B., Zamora, S. & Álvaro, J. J. 2013. The oldest echinoderm faunas from Gondwana show that echinoderm body plan diversification was rapid. Nature Communications, 4: 1385 doi: 10.1038/ncomms2391.
- Zamora, S., Rahman, I. A. & Smith, A. B. 2013. The ontogeny of cinctans (stem-group echinodermata) as revealed by a new genus, Graciacystis, from the middle Cambrian of Spain. Palaeontology 56: 399-410.
- Zamora, S. 2013. Morphology and phylogenetic interpretation of a new cambrian edrioasteroid (Echinodermata) from Spain. Palaeontology 56: 421-431.
- Zamora, S., Darroch, S. & Rahman, I. A. 2013. Taphonomy and ontogeny of early pelmatozoan echinoderms: a case study of a mass mortality assemblage of Gogia from the Cambrian of North America. Palaeogeography, Palaeoclimatology, Palaeoecology. 377, 62-72.
- Zamora, S., Sumrall, C. D. & Vizcaïno, D. 2013. Morphology and ontogeny of the Cambrian edrioasteroid (Echinodermata) Cambraster cannati (Miquel) from western Gondwana. Acta Palaeontologica Polonica, 58 (3): 545-559.
  Zamora, S., Lefebvre, B., Álvaro, J. J., Clausen, S., Elicki, O., Fatka, O., Jell, P., Kouchinsky, A., Lin, J-P., Nardin, E., Parsley, R., Rozhnov, S., Sprinkle, J., Sumrall, C. D., Vizcaïno, D. & Smith, A. B. 2013. Cambrian echinoderm diversity and palaeobiogeography. From: Harper, D. A. T. & Servais, T. (eds) 2013. Early Palaeozoic Biogeography and Palaeogeography.. Geological Society of London, Memoirs. 38, 157-171.


- Zamora, S. & Smith, A. B. 2012. Cambrian stalked echinoderms show unexpected plasticity of arm construction. Proceedings of The Royal Society B, 279: 293-298.
- Zamora, S. 2012. The first Furongian (late Cambrian) echinoderm from the British Isles. Geological Magazine, 149, 940-943.
- Zamora, S., Rahman, I. A. & Smith, A. B. 2012. Plated Cambrian bilaterians reveal the earliest stages of echinoderm evolution, PLoS One 7(6).
- Esteve, J. Sundberg, F., Zamora, S. & Gozalo, R. A new Alokistocaridae Resser, 1939 (Trilobita) from the Middle Cambrian of Spain. Geobios, 45, 275-283.
- Mergl, M. & Zamora, S. New and revised occurrences of rhynchonelliformean brachiopods from the middle Cambrian of the Iberian Chains, NE Spain. Bulletin of Geosciences, 87 (3), 571-586.


- Sumrall C. D. and Zamora S. 2011. Ordovician Edrioasteroids from Morocco: Faunal exchanges across the Rheic Ocean. Journal of Systematic Palaeontology, 9(3): 425-454.
- Zamora, S. 2011. Equinodermos del Cámbrico de España: Situación actual de las investigaciones y perspectivas futuras. Estudios Geológicos, 67(1): 59-81.
- Esteve, J., Hughes, N. & Zamora, S. 2011. The Purujosa trilobite assemblage and the evolution of trilobite enrollment. Geology, 39 (6): 575-578.
- Zamora, S., Mayoral, E., Esteve, J., Gámez J. A. & Santos, A. 2011. Skeletal abnormalities in paradoxidid trilobites from the Cambrian of Spain. Bulletin of Geosciences, 86 (3): 665-673.


- Zamora, S. 2010. Middle Cambrian echinoderms from North Spain show echinoderms diversified earlier in Gondwana. Geology, 38: 507-510.
- Rahman, I.A., Zamora, S. & Geyer, G. 2010. The oldest stylophoran echinoderm: a new Ceratocystis from the Middle Cambrian of Germany. Paläontologische Zeitschrift, 84 (2): 227-237.
- Zamora, S. & Smith, A. B. 2010. The oldest isorophid edrioasteroid (Echinodermata) and the evolution of attachment strategies in Cambrian edrioasteroids. Acta Palaeontologica Polonica, 53 (3):487-494.
- Zamora, S. & Álvaro, J.J. 2010. Testing for a decline in diversity prior to extintion: Languedocian (latest mid-Cambrian) distribution of cinctans (Echinodermata) in the Iberian Chains, NE Spain. Palaeontology, 53 (6): 1349-1368.
- Esteve, J.V., Zamora, S., Gozalo, R. & Liñán, E. 2010. Sphaeroidal enrollment in middle Cambrian solenopleuropsine trilobites. Lethaia, 43 (4): 478-493.
- Zamora, S., Clausen, S., Álvaro, J.J. & Smith, A.B. 2010. Pelmatozoan echinoderms colonized carbonate firm ground substrates in hight energy environments since the basal Middle Cambrian. Palaios, 25 (12): 764-768.


- Smith, A.B. & Zamora, S. 2009. Rooting phylogenies of problematic fossil taxa; a case study using cinctans (stem-group echinoderms). Palaeontology, 52 (4): 803-821.
- Zamora, S., Álvaro J. J. & Vizcaïno., 2009. Pelmatozoan echinoderms from the Cambrian-Ordovician transition of the Iberian Chains (NE Spain): early diversification of anchoring strategies. Swiss Journal of Geosciences, 102: 43-55.
- Zamora, S., Gozalo, R. & Liñán E., 2009. Middle Cambrian gogiid echinoderms from Northeast Spain: Taxonomy, palaeoecology and palaeogeographic implications. Acta Paleontologica Polonica, 54(2): 253-265.
- Rahman, I.A. & Zamora, S., 2009. The oldest cinctan carpoid (stem-group Echinodermata) and the evolution of the water vascular system. Zoological journal of the Linnean Society, 157: 420-432.


- Zamora, S., Mayoral, E., Gámez Vintaned, J. A., Bajo, S. & Espílez, E. 2008. The infaunal echinoid Micraster: taphonomic pathways indicated by sclerozoan trace and body fossils from the Upper Cretaceous of northern Spain. Geobios, 41: 15-29.
- Zamora, S. & Smith, A. B. 2008. A new Middle Cambrian stem group echinoderm from Spain: Palaeobiological implications of a highly asymmetric cinctan.  Acta Palaeontologica Polonica, 53(2): 207-220. Most Cited Acta Palaeontologica Paper. Honorable Mention for 2009.
- Liñán, E., Díes M. E., Gozalo R. Gámez Vintaned J. A., & Zamora, S., 2008. Nuevos trilobites del Ovetiense inferior (Cámbrico Inferior bajo) de Sierra Morena (España) Ameghiniana, 45(1): 123-138.


- Zamora, S., Liñán, E., Domínguez Alonso, P., Gozalo, R. & Gámez Vintaned, J. A. 2007. A Middle Cambrian edrioasteroid from the Murero biota (NE Spain) with Australian affinities. Annales de Paleontologie, 93: 249-260.
  Zamora, S., Liñán, E., Gámez Vintaned, J. A., Domínguez Alonso, P. & Gozalo, R. 2007. Nuevo carpoideo de la Clase Cincta Jaeckel, 1918 del norte de España: Inferencias sobre la morfología funcional del opérculo. Ameghiniana, 44-4: 227-238.
Foto:Samuel Zamora Iranzo

Panoramic view of Cambrian rocks from the Anti-Atlas (Morocco)
Foto:Samuel Zamora Iranzo

Echinoderm fossils from the Spanish Paleozoic

Imagen Boceto IGME


Ríos Rosas, 23

28003 Madrid

Teléfono + 34 913 495 700

Icono de conformidad con el Nivel A, de las Directrices de Accesibilidad para el Contenido Web 1.0 del W3C-WAI