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Enlaces de interés

Dr. Enrique Peñalver Mollá

Enrique Peñalver Mollá
Current position:
Scientific Researcher, Geological Survey of Spain (Instituto Geológico y Minero de España – IGME)

Previous positions/career:
- Researcher Contract “Ramón y Cajal” (since 2007 to 2009), Spanish Ministry and Museo Geominero (IGME).
- Post Doctoral Fellow (since February 2004 to February 2006) in the American Museum of Natural History (USA).
- PhD (concluded 2002), University of Valencia – “Los insectos dípteros del Mioceno del Este de la Península Ibérica: Rubielos de Mora, Ribesalbes y Bicorp. Tafonomía y sistemática”.

Phone: 00 34 91 3495939
Email: e.penalver@igme.es

Mailing address:
Museo Geominero
Instituto Geológico y Minero de España
c/ Ríos Rosas 23
28003 Madrid, Spain

Scientific research:
My main interests in paleoentomology are taxonomy, taphonomy and paleoecology (paleobiology). I study compression fossils (mainly Miocene in age) and bioinclusions in amber (mainly Cretaceous in age) and copal (Quaternary). Practically all my research is focused in insects, but a few of my publications are on arachnids and crustaceans. I am very interested in search of new paleontological outcrops in Spain. Key words about my research are: Miocene, Cretaceous, Albian, Dominican amber, Spanish amber (El Soplao, San Just and Álava ambers), New Zealand copal, Spanish compression fossils (Rubielos de Mora, Ribesalbes and Bicorp outcrops), paleolakes, rithmites (oil shales), insects, Hymenoptera, Orthoptera, Diptera, Lepidoptera, Hemiptera, Coleoptera, Thysanoptera, Arachnida, ticks, Opiliones, Crustacea, Tanaidacea, taphonomy, microbial mats, disarticulation, exceptional preservation, the amber origin, amphibians, paleoecology, paleobiology, plant-insect interactions, early insect pollination, early camouflage in insects, vertebrate-arthropod interactions, hematophagy, litter microhabitat, Tertiary insect swarming, past reproductive behaviours.

Doctoral Thesis and post-doctoral research:
My PhD research assessed the paleodiversity of the order Diptera in the Miocene lacustrine sediments of Rubielos de Mora, Ribesalbes and Bicorp (Spain), with special attention to taphonomy and paleoecology: In the Taphonomy section actualistic experiments were accomplished to know the processes of flotation, collapse, decomposition and disarticulation of terrestrial Diptera in lakes, and the results have been applied to the interpretation of the fossil record. The term syncompression was proposed for a more methodical analysis of the taphonomy and palaeoecology in the outcrops of compression. It was deduced that microbial mats covered the bottom of the palaeolakes studied, and that these mats played a very important role in the fossilization of the insects. In the Systematic section the different Diptera’s taxa found were studied and grouped in 18 families and 28 genera. The Palaeoecology section was focused on three questions: evidence of trophic behavior, evidence of Chironomidae and Acrididae swarms, and the study of egg depositions on leaves by Odonata. A reconstruction attempt of the limnic food web for the three paleolakes was accomplished.”

My post-doctoral research was conducted since February 2004 to February 2006 in the American Museum of Natural History in New York, under the direction of Dr. David Grimaldi. I studied selected Dominican amber pieces housed at the AMNH collection with uncommon fossil assemblages of special palaeoecological interest. Some of the topics studied were fig pollination by fig wasps, assemblages of mammalian hair and blood-feeding midges (hematophagy) and butterflies related to palaeobiogeographic interesting data and the age of origin of butterflies.

Current tasks:
Currently I am a Scientific Researcher in the Museo Geominero (IGME) in Madrid. My research mainly is focused on insects preserved in Albian amber of Spain, under the project CGL2014-52163 of the Spanish Ministry of Economy and Competitiveness. Currently, I am especially interested in Cretaceous ticks, the systematics of the Order Thysanoptera, on taphonomic studies of the Hymenaea resin in Madagascar, and diverse aspects of the copal from New Zealand.


SCI articles (last decade):

Peñalver, E.; Grimaldi, D.A. & Delclòs, X. 2006. Early Cretaceous Spider Web with Its Prey. Science, 312: p. 1761.

Engel, M.S. & Peñalver, E. 2006. A Miocene Halictine Bee from Rubielos de Mora Basin, Spain (Hymenoptera: Halictidae). American Museum Novitates, 3503: 10 pp.
I.S.S.N. 0003-0082

Peñalver, E. & Grimaldi, D. 2006. Assemblages of mammalian hair and blood-feeding midges (Insecta: Diptera: Psychodidae: Phlebotominae) in Miocene amber. Transactions of the Royal Society of Edinburgh, Earth Sciences, 96: 177-195.

McNamara, M.; Orr, P.J.; Kearns, S.L., Alcalá, L.; Anadón, P. & Peñalver, E. 2006. High-fidelity organic preservation of bone marrow in ca.10 Ma amphibians. Geology, 34 (8): 641-644.

Peñalver, E. & Grimaldi, D. 2006. New data on Miocene amber butterflies in Dominican amber (Lepidoptera: Riodinidae and Nymphalidae) with the description of a new nymphalid. American Museum Novitates, 3519: 17 pp.

Peñalver, E.; Engel, M. & Grimaldi, D.A. 2006. Fig Wasps in Dominican Amber (Hymenoptera: Agaonidae). American Museum Novitates, 3541: 16 pp.

Peñalver, E. & Engel, M. 2006. Two Wasp Families Rare in the Fossil Record (Hymenoptera): Perilampidae and Megaspilidae from the Miocene of Spain. American Museum Novitates, 3540: 12 pp.

Delclòs, X.; Arillo, A.; Peñalver, E.; Barrón, E.; Soriano, C.; López del Valle, R.; Bernárdez, E.; Corral, C. & Ortuño, V.M. 2007. Fossiliferous amber deposits from the Cretaceous (Albian) of Spain. Comptes Rendus Palevol, 6: 135-149.

Peñalver, E.; Álvarez-Fernández, E.; Arias, P.; Delclòs, X. & Ontañón, R. 2007. Local amber in a Palaeolithic Context in Cantabrian Spain: the case of La Garma A. Journal of Archaeological Science, 34: 843-849.

Peñalver, E.; Delclòs, X. & Soriano, C. 2007. A new rich amber outcrop with palaeobiological inclusions in the Lower Cretaceous of Spain. Cretaceous Research, 28: 791-802.

Corchón, M.S., Mateos, A., Álvarez-Fernández, E., Peñalver, E., Delclòs, X. & Van der Made, J. 2008. Ressources complémentaires et mobilité dans le Magdalénien cantabrique. Nouvelles données sur les mammifères marins, les crustacés, les mollusques et les roches organogènes de la Grotte de Las Caldas (Asturies, Espagne). L´Anthropologie, 112: 284-327.

Arillo, A.; Peñalver, E. & Delclòs, X. 2008. Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits. Zootaxa, 1920: 29-40.

McNamara, M.E.; Orr, P.J.; Kearns, S.T.; Alcalá, L.; Anadón, P. & Peñalver, E. 2009. Soft-tissue preservation in Miocene frogs from Libros, Spain: insights into the genesis of decay microenvironments. Palaios, 24: 104-117.

Arillo, A.; Peñalver, E. & García-Gimeno, V. 2009. First fossil Litoleptis (Diptera: Spaniidae) from the Lower Cretaceous amber of San Just (Teruel Province, Spain). Zootaxa, 2026: 33-39.

Najarro, M.; Peñalver, E.; Rosales, I.; Pérez-de la Fuente, R.; Daviero-Gomez, V.; Gomez, B. & Delclòs, X. 2009. Unusual concentration of Early Albian arthropod-bearing amber in the Basque-Cantabrian Basin (El Soplao, Cantabria, Northern Spain): Palaeoenvironmental and palaeobiological implications. Geologica Acta, 7 (3): 363-387.

McNamara, M.; Orr, P.J.; Kearns, S.T.; Alcalá, L.; Anadón, P. & Peñalver, E. 2009. Exceptionally preserved tadpoles from the Miocene of Libros, Spain: ecomorphological reconstruction and the impact of ontogeny upon taphonomy. Lethaia, 43: 290-306. DOI 10.1111/j.1502-3931.2009.00192.x

McNamara, M.; Orr, P.J.; Kearns, S.T.; Alcalá, L.; Anadón, P. & Peñalver, E. 2009. Organic preservation of fossil musculature with ultracellular detail. Proceedings of the Royal Society, Series B, 277: 423-427.

Pérez-de la Fuente, R.; Nel, N.; Peñalver, E. & Xavier Delclòs, X. 2010. A new Early Cretaceous snakefly (Raphidioptera: Mesoraphidiidae) from El Soplao amber (Spain). Annales de la Société entomologique de France, 46 (1-2): 108-115.

Peñalver, E. & Nel, P. 2010. Hispanothrips from Early Cretaceous Spanish amber, a new genus of the resurrected family Stenurothripidae (Insecta: Thysanoptera). Annales de la Société entomologique de France, 46 (1-2): 138-147.

Nel, P.; Peñalver, E., Azar, D., Hodebert, G. & Nel, A. 2010. Modern thrips families Thripidae and Phlaeothripidae in the Early Cretaceous amber (Insecta: Thysanoptera). Annales de la Société entomologique de France, 46 (1-2): 154-163.

Peñalver, E. & Grimaldi, D. 2010. Latest occurrences of the Mesozoic family Elcanidae (Insecta: Orthoptera), in Cretaceous amber from Myanmar and Spain. Annales de la Société entomologique de France, 46 (1-2): 88-99.

Peñalver, E.; Ortega, J.; Nel, A. & Delclòs, X. 2010. Mesozoic Evaniidae (Insecta: Hymenoptera) in Spanish Amber: Reanalysis of the Phylogeny of the Evanioidea. Acta Geologica Sinica (English Edition), 84 (4): 809-827.

Najarro, M.; Peñalver, E.; Pérez-de la Fuente, R.; Ortega-Blanco, J.; Menor-Salván, C.; Barrón, E.; Soriano, C.; Rosales, I.; López del Valle, R.; Velasco, F.; Tornos, F.; Daviero-Gomez, V.; Gomez, B. & Delclòs, X. 2010. Review of the El Soplao amber outcrop, Early Cretaceous of Cantabria, Spain. Acta Geologica Sinica (English Edition), 84 (4): 959-976.

Peñalver, E. & Gaudant, J. 2010. Limnic food web and salinity of the Upper Miocene Bicorb palaeolake (eastern Spain). Palaeogeogr., Palaeoclimatol., Palaeoecol., 297: 683-696.

Ortega, J.; Delclòs, X.; Peñalver, E. & Engel, M.S. 2011. Serphitid wasps in Early Cretaceous amber from Spain (Hymenoptera: Serphitidae). Cretaceous Research, 32: 143-154.

Ortega, J.; Peñalver, E.; Delclòs, X. & Engel, M.S. 2011. False fairy wasps in Early Cretaceous amber from Spain (Hymenoptera: Mymarommatoidea). Palaeontology, 54 (3): 511-523.

Pérez-de la Fuente, R., Delclòs, X, Peñalver, E. & Arillo, A. 2011. Biting midges (Diptera: Ceratopogonidae) from the Early Cretaceous El Soplao amber (N Spain). Cretaceous Research, 32: 750-761.

McNamara, M.; Orr, P.J.; Manzocchi, T.; Kearns, S.T.; Alcalá, L.; Anadón, P. & Peñalver, E. 2011. Biological controls upon the physical taphonomy of exceptionally preserved salamanders from the Miocene of Rubielos de Mora, Spain. Lethaia: 17 pp. DOI: 10.1111 ⁄j.1502-3931.2011.00274.x

McNamara, M.; Orr, P.J.; Alcalá, L.; Anadón, P. & Peñalver, E. 2012. What controls the taphonomy of exceptionally preserved taxa- environment or biology? A case study using frogs from the Miocene Libros Konservat-Lagerstätte (Teruel, Spain). PALAIOS, 27: 63-77.

Peñalver, E.; Labandeira, C.C.; Barrón, E.; Delclòs, X.; Nel, P.; Nel, A.; Tafforeau, P. & Soriano, C. 2012. Thrips Pollination of Mesozoic Gymnosperms. PNAS, 109 (22): 8623-8628.

Pérez-de la Fuente, R.; Peñalver, E.; Delclòs, X. & Engel, M.S. 2012. Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida: Raphidioptera). ZooKeys, 204: 1-40.

Pérez-de la Fuente, R.; Peñalver, E. & Ortega-Blanco, J. 2012. A new species of the diverse Cretaceous genus Cretevania Rasnitsyn, 1975 (Hymenoptera: Evaniidae) from Spanish amber. Zootaxa, 3514: 70-78.

Pérez-de la Fuente, R.; Delclòs, X.; Peñalver, E.; Speranza, M.; Wierzchos, J.; Ascaso, C.; Engel, M.S. 2012. Early evolution and ecology of camouflage in insects. PNAS, 190 (52): 21414-21419.

Peñalver, E.; Fontal-Cazalla, F.M. & Pujade-Villar, J. 2013. Palaeogronotoma nov. gen. from the Miocene of Spain, the first Tertiary fossil record of the subfamily Eucoilinae (Hymenoptera: Figitidae). Geodiversitas, 35 (3): 643-653.

Peñalver, E. & Pérez-de la Fuente, R. 2014. Unearthing the secrets of ancient immature insects. eLife: 3 pp. DOI: 10.7554/eLife.03443

Sánchez-García, A.; Peñalver, E.; Pérez-de la Fuente, R. & Delclòs, X. 2015. A rich and diverse tanaidomorphan (Crustacea: Tanaidacea) assemblage associated with Early Cretaceous resin-producing forests in North Iberia: palaeobiological implications. Journal of Systematic Palaeontology, 13 (8): 645-676.

Arillo, A.; Peñalver, E.; Pérez-de la Fuente, R.; Delclòs, X.; Criscione, J.; Barden, P. M.; Riccio, M. L. & Grimaldi, D. A. 2015. Long-proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae). Systematic Entomology, 40: 242-267.

Peris, D.; Solórzano Kraemer, M.M.; Peñalver, E. & Delclòs, X. 2015. New ambrosia beetles (Coleoptera: Curculionidae: Platypodinae) from Miocene Mexican and Dominican ambers and their paleobiogeographical implications. Organisms Diversity and Evolution, 15: 527-542. DOI: 10.1007/s13127-015-0213-y

Gaudant, J.; Barrón, E.; Anadón, P.; Reichenbacher, B. & Peñalver, E. 2015. Palaeoenvironmental analysis of the Miocene Arcas del Villar gypsum sequence (Spain), based on palynomorphs and cyprinidontiform fishes. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 277 (1): 105-124.

Peñalver, E.; Arillo, A.; Riccio, M.L.; Pérez-de la Fuente, R.; Delclòs, X.; Barrón, E. & Grimaldi, D.A. 2015. Long-proboscid Flies as Pollinators of Cretaceous Gymnosperms. Current Biology, 25 (14): 1917-1923.

Speranza, M.; Ascaso, C.; Delclòs, X. & Peñalver, E. 2015. Cretaceous mycelia preserving fungal polysaccharides: Taphonomic and paleoecological potential of microorganisms preserved in fossil resins. Geologica Acta, 13 (4): 409-431.

Pérez-de la Fuente, R.; Delclòs, X.; Peñalver, E.; Engel, M.S. 2016. A defensive behavior and plant-insect interaction in Cretaceous amber ‒ the case of Hallucinochrysa diogenesi. Arthropod Structure and Development, 45: 133-139.

Delclòs, X.; Peñalver, E.; Arillo, A.; Engel, M.S.; Nel, A.; Azar, D. & Ross, A. 2016. New mantises (Insecta: Mantodea) in Cretaceous ambers from Lebanon, Spain and Myanmar. Cretaceous Research, 60: 91-108.

McNamara, M.E.; Orr, P.J.; Kearns, S.; Alcalá, L.; Anadón, P. & Peñalver, E. 2016. Reconstructing carotenoid-based and structural coloration in fossil skin. Current Biology, 26: 1075-1082.

Books, scientific comics and DVDs of popularization of Science:

Peñalver, E. 1998. Estudio tafonómico y paleoecológico de los insectos del Mioceno de Rubielos de Mora (Teruel). Instituto de Estudios Turolenses Ed.: 179 pp. Teruel.

Peñalver, E. 2001. Los insectos fósiles de Rubielos de Mora. Excmo. Ayuntamiento de Rubielos de Mora Ed., 31 pp. Teruel.

Peñalver, E. 2002. Los insectos dípteros del Mioceno del Este de la Península Ibérica; Rubielos de Mora, Ribesalbes y Bicorp. Tafonomía y sistemática. Tesis Doctoral. Universitat de València: 548 pp. Valencia. (formato PDF en CD-Rom).

Belinchón, M.; Peñalver, E.; Montoya, P. y Gascó, F. 2009. Crónicas de fósiles. Las colecciones paleontológicas del Museo de Ciencias Naturales de Valencia. Ayuntamiento de Valencia: 538 pp.

Rodrigo, A.; Peñalver, E.; Barrón, E. y Lozano, R.P. 2010. Gea y el ámbar. Instituto Geológico y Minero de España, Madrid: 27’

Peñalver, E. 2012. Ámbar. La historia de un material que ha fascinado a la humanidad. Los Libros de la Catarata: 134 pp. Madrid.

Rodrigo, A.; Peñalver, E.; Barrón, E. y Lozano, R.P. 2015. Gea y los fósiles/Gea and fossils. Instituto Geológico y Minero de España, Madrid: 28’

Solórzano-Kraemer, M.M.; Delclòs, X.; Peñalver, E. & Rodrigo, A. 2016. Vongy. An Adventure Among Scientists/Vongy. Une aventure entre scientifiques/ Vongy. Ein Abenteuer unter Wissenschaftlern/ Vongy. Una aventura entre científicos. National Geographic-Spanish Ministry of Economy and Competitiveness- German Research Foundation: 28 pp.

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